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As it is written in the relic screen:

Below is the current status for the relic Unholy Shrine. By capturing and holding a relic, your guild members will received the bonuses detailed below until control of the relic is lost. Be sure to keep at least one guild member to guard it at all times (note however other guilds may use groups to attempt to capture it).

The lead defender (highest level) receives a 50% bonus to their stats when attacked (providing you do not control any other relics - note also if you control 3 or more relics, the lead defender will suffer a 10% reduction to their stats at each relic).

Each additional defending guild member will apply 20% of there stats when calculating the total defense (in the same way as a group attack) providing they have been active (logged in) within the last 7 days